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A windshirt is cool, comfortable and quick-drying enough to wear as a lone base layer. If you are exerting yourself in a mild, damp climate, and need to keep yourself covered for reasons of a) decency, and b) fending off small, biting flies, and disease-carrying ticks, it is the best option. Whilst Merino is nice to wear, a wool base layer does not take well to scrambling amongst rocks and brush, and will not dry as fast as a windshirt if you get soaked. I find that as long as I’m moving uphill, a windshirt alone is enough down to about -5 degrees C. If you are walking up a mountain in slushy snow, and need to pause, you can throw a pertex & pile top or Paramo over the windshirt, without feeling clammy underneath. If you have garments that dry fast, you don’t worry so much about getting wet, and you don’t feel like a boil-in-the-bag sweaty mess.
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